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All Bosses in Mortal Shell 2

Every boss, where it waits, and what beating it pays out. Each one gates something — a Shell, a Seal, a lever — so this doubles as the order the game expects you to take them in.

45catalogued
103things they drop
40/45with art

How boss fights work

A boss is not a health bar you outlast. Health and Break run as two separate meters, and the second is the one that ends fights: land your Seal's defence at the right moment, fill the Break meter, and the opening you get is a Riposte that costs the boss more than a minute of chip damage would.

Almost every boss gates something — a Shell, a Seal, a lever, a weapon — so the drops list is also the progression map. If you are stuck on where to go next, the boss you have not beaten is usually holding it.

Learn the Seal, not the bossGuard, Parry and Harden each have a different window. Most fights that feel impossible are a timing you have not settled on yet.
The ending is in this listFive of these are endgame bosses and every one is tagged. Nothing about the last hours of the game is stated in a row you did not open.
Some fights are scriptedAt least one boss ends in a cutscene whatever you do. Losing is not always losing.
Bloodcursed LithopodGloomshade Grove, FainwealdCitadel Gate Lever, Coins, Gloom
Bor, the Deluded MonarchCleanse the Abbey Entrance BeaconGlimpse, Monarch's Vestige, Gloom
Caged GrishaCleanse the Winterglass Lake Beacon in the Conquered TempleGrisha Remnant, Fusillade Stone, Coin, Gloom
Circle of the Grasping RootRootwater Hollow, through the Seeping Mire beaconGloom, Coin
Droeg, the ConquererConquered TempleConqueror's Reward, Gloom
Gloombound LegionCleanse the Hangman's Scaffold Beacon in the Prisoner's Domain GateGloom
Gloombound RitualistSunken ground past the Village Outskirts beaconGloom
Gloombound WraithForbidden ArchivesGlimpse, x2, Gloom, Curseblood Stone
Great Arbiter of FleshWidow's OverlookGlimpse x1, Gloom, Arbiter's Prize
GrishaFound deep inside the Flooded Village dungeon past the small arena.Gloom, Synaptic Vessel, Grisha Remnant
HexapodWithered ShoalsHexapod Core, Gloom
Infested MinerBurrowkin Dwelling, Ruins of MammonGlimpse, Ventrium, Laterite, Dorsalite, Wretchcaller's Stone, Coins, Gloom
Isaac, The Scholar-PrinceAshen SanctumScholar's Wormstone, Gloom
Lucian, the Thirsting KnightThe Hidden KeepSerpent Stone, Glimpse, 6500 Gloom
Magdalena the Lady of the WoodsSunken VillageGloom, Magdalena's Memento
Malborn OffspringendgameThe tower it patrols, beneath Marrow Keep
Nameless CaptivePrisoner's DomainCaptive's Scabstone, Gloom
Orrem, the Discarded GolemendgameBeneath Marrow Keep, guarding one of the three infection points
Pale GrishaFound in the Grisha Hunting GroundsRevered Gland, Grisha Remnant, Gloom
Prophet of Profane InfinitiesObsidianite MinesObsidian Hammer, Paradoxical Scripture, Coins, Gloom
Rabid Grisha PackFound at the bottom of the Holding Cells dungeonGrisha Remnant, Inflamed Clawstone, Gloom
Sariel, the EndlessInitially found on the platform north of the Chamber of Becoming and moves inside the chamber during 2nd phase.Gloom, Glimpse, Clockwork Scythe, Sariel the Endless
Servants of the Scholar PrinceCleanse the Citadel Atrium BeaconGloom
Sester SecundusRevenant GravesSester's Censer, Gloom
Solnir, the Gloom SeekerCleanse the Faded Citadel Gate BeaconGlimpse, Solnir Shard, Gloom
Starved HarridanFound south of the Castigator's Keep BeaconVoltaic Essence, Voltaic Amber, Gloom
Strange GrishaFound in the Withered ShoalsGrisha Remnant, Strange Remnant, Gloom
Stray SestersCleanse the Sester's Gate BeaconBlack Needle, Sester's Crest, Gloom
Subjugated GuardianSentry's GraveBallistazooka, Coins, Gloom
Tainted VestigeHidden cave off the Disciple's GrottoGloom
Tar GolemDisciple's GrottoVatra's Seal
Tarblighted StonerSanguine CavernsTarred Fragment, Gloom
The Foul PassengerendgamePulled out of Zmey, the Unbidden
The Lost ChildSanguine Caverns (Gate)Lost Clotstone, Gloom
The MonolithendgameBeneath Marrow Keep, guarding one of the three infection points
The Wandering ShepherdFarm compound across the bridge from the Sunken Village beaconTarblighted Trophy, Gloom
The WardenCitadel Of PenanceGloom, access to claim Eredrim as a shell
Tishina's ConfessorFound at the end of Slumwater Drain in the Prisoners' Domain region.Confessor's Keepsake, Gloom
Twin SestersSester's BastionBlack Needle, Sester's Crest, Gloom
Urrig, the ExecutionerFound in the Conquered TempleWarden's Stone, Gloom
Vellen, High Lord of MammonRoyal Crypt of MammonGlimpse, Gloom
Viletongue BatushkaCleanse the Castigator's Keep BeaconGlimpse, Sage's Gate Lever, Viletongue Hedron
Vrago, Solikar ChampionAbandoned ShrineVoltaic Crown, Gloom
Vrannic the Grand IllusionistHall of Illusions, through the Mushroom Village Gate beaconVolatile Fragment, Glimpse, Gloom
Zmey, the UnbiddenendgameBeneath Marrow Keep, once all three branches are severed